In Anânh, Brood Queen
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In Anânh, Brood Queen is a large, high-ranking Hive Wizard who is the consort of Hiraks, the Mindbender, as well as the source of Hive on the Tangled Shore. She is the main boss of the Broodhold Strike.
In Anânh's most dangerous ability lies in being able to change into a shade form, switching from a Solar to Void power-set, both of which share the Burning Debuff on all attacks that causes damage over time. When first spawned, In Anânh is Solar-based and has the ability to fire searing Solar fireballs that explode at Guardians and an explosive Solar melee that acts like a Ground Pound but with Solar damage. She also has the spell "Fire Nova Gaze", where she charges Solar energy in her hands that also causes Burning to all Guardians and, when the countdown of 3 seconds on the left of the HUD reaches zero, launches an explosive fiery wave called "Nova Scream" across the room that will wipe out Guardians if they are not at full health, in a Healing Rift, or hiding behind the rocks in the area which will prevent damage.
At two-thirds health, she will reach out to Ur, the Ever-Hunger and create a Shade of herself with Void-based abilities. The Shade fires Void blasts more rapidly, but they are not explosive, as well as having the same melee but with Void damage. Her main ability is to set down multiple Void traps called "Brood Spikes", which do damage over time when a Guardian stands in one through Burning, before they explode after a couple seconds and launching guardians that survive into the air.
Upon reaching the last third of her health, the two forms become two separate entities entirely with both using their respective abilities at once, forcing Guardians to deal with two Strike bosses instead of one, adding a much greater threat into the mix. When one dies, the remaining one will become enraged and enter a Berserk Mode.