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Uldren Sov, born Uldwyn Sov as a human, was one of the first Awoken and is the loyal brother of the Queen of the Reef Mara Sov. As the Prince of the Reef he served as the Master of Crows, the official spymaster of the Reef and monitored all threats to his people. While a loyal advisor to his sister, he disagreed with Mara frequently on matters of defense and foreign policy, and was very disdainful of the Guardians. During the Battle of Saturn his fighter was damaged and crashed on Mars. Over the next several years Uldren sought to find his sister, convinced she remained alive, but was corrupted and manipulated by Riven into creating the Scorn and turning on the Reef in his bid to reopen the Dreaming City. After murdering Cayde-6 during his crusade, Uldren was eventually hunted down and killed by the Young Wolf and Petra Venj.
Uldren's body was found in the Dreaming City by a Ghost and revived as a Lightbearer. Traveling to Earth and naming his Ghost Glint, the new Lightbearer soon learned to hide from Guardians who grew violent when they saw his face. After one confrontation left him drifting in the vacuum of space, he was rescued by The Spider, who mockingly dubbed him as The Crow and pressed him into service as an enforcer to repay his debt. Crow was freed from this servitude by the Young Wolf after helping them defeat the High Celebrant of Xivu Arath and chose to become a Guardian after receiving visions from the Traveler. Under the mentorship of Osiris, Uldren traveled to the The Last City and began working as a scout for the Vanguard.
Uldren Sov's Biography
During the Golden Age, the man who would become Uldren Sov was Uldwyn, one of the nine hundred crewmembers of the colony ship Yang Liwei. He served aboard the ship as part of Project Amrita along with his mother Osana and his fraternal twin sister Mara. He was known to be boisterous and a risk-taker, often prize-fighting with other members of the crew.
As the Yang Liwei was outbound from the Solar System, the Darkness descended to bring about the Collapse. As the Yang Liwei was engulfed by the black, light-negating field broadcast by the Darkness, the Traveler struck out with a beam of Light and held the Darkness at bay. The force of the conflict between the two paracausal forces was too great, and a kugelblitz formed, swallowing the Yang Liwei and all of its passengers and crew, including Uldwyn. 
Rebirth as an Awoken
Like the rest of the Awoken, Uldwyn was reborn in the pocket universe known as the Distributary with little to no memory of his previous life. He took the name Uldren, which like most Awoken names is a permutation of his original human name.
Breaching the Black Garden
Long before the resurrection of the Guardian, Uldren was beloved and admired by the Awoken people. He was famous for many deeds, but he knew underneath it all, the love of the people was due to his association with Mara. Seeking to be more than that to others, and to do something that even his seemingly all-knowing sister couldn't anticipate, he resolved to venture into the Black Garden with only his wing-man Jolyon. The duo ventured into Meridian Bay, fending off the Cabal and Vex as they search for a way to trick the latter race into opening the gate to the Garden. Eventually, Uldren has a master plan: lure out a Gate Lord to get the gate open, then agitate the trigger-happy Cabal into killing the Gate Lord. The plan worked, and the two spent an undetermined amount of time exploring the Black Garden. During their venture, Uldren found beauty in the Garden. He developed a desire to protect it from the Guardians and a desire to one day bring Mara there. During his journey, he also attacked a detachment of Cabal that had been pulled into the Garden by the Vex, and learned that the Garden's secrets can "grow" into one's body and mind from an injured Legionary.
Uldren and Joylon eventually retreated from the Garden and returned to the Reef, battered in body and mind by their adventure. He returned to Mara and presented a clump of asphodelia he had picked in the Garden. Mara was initially impressed, but once the two were alone, Mara asked Uldren about the Garden's heart, then expressed displeasure that he ignored her warning to not go to the Garden. While thinking about the Heart, he realized that it may be bait to lure the Guardians to the Garden and mark a milestone in the Traveler's recovery, and that he still failed to surprise his sister. The latter thought was the one that wounded him the most. Later, he talked to Joylon, who asks Uldren not to make what he saw in the Garden public.
The Reef Wars
Uldren intercepted a Fallen transmission that was broadcast on all channels but was unable to decrypt it, and was only able to discern that all of the Fallen Houses were having an unprecedented level of communication and cooperation. The Techeuns looked toward Earth and saw the Fallen gathering there, and the Reef's leadership came to the conclusion that the Fallen were joining forces for a major attack on the Last City. At first, they believed there was nothing they could do, until the House of Wolves passed through the asteroid belt and stopped at Ceres. There, the Awoken revealed themselves and assaulted the Wolves, using the Harbingers to shatter the planetoid and killed half the House of Wolves, including Virixas, Kell of Wolves. However, the Battle of Ceres was only the start of the Reef Wars, as the remaining members of the House of Wolves scattered throughout the Reef and split into three factions, led by Irxis, Wolf Baroness; Parixas, the Howling; and Skolas, the Rabid.
After two major losses to the Wolves at Amethyst and Hygiea and a pyrrhic victory at Iris, Uldren provided the Awoken with a major advantage when he and his Crows cracked the Fallen's encryptions. Using knowledge gained from the intercepted communications, Uldren assisted Paladin Imogen Rife in luring one of Skolas' vassals, Baroness Drevis, into a trap at the asteroid Bamberga which destroyed her Ketch and led to the capture of Drevis, Kaliks-4, and many other high-ranking Fallen. Unfortunately, the victory was short-lived as Rife's fleet was ambushed by another of Skolas' vassals, Pirsis, who gathered a massive fleet and laid siege to the settlement of Pallas, trapping Rife's forces for years. The stalemate was broken when Weksis, the Meek launched a foolhardy assault on Pallas that failed, giving the Reef superior numbers on the Fallen. Uldren and his Crows then worked with the Techeuns to create an illusion of the Harbingers, which scared the Wolves into scattering and led to the capture of Pirsis, although the Fallen captain had managed to slay Paladin Rife during the battle.
Following the capture of Beltrik, the Veiled at Fortuna Plummet, Uldren was brought a message by one of his Crows from a Fallen named Variks, the last member of the House of Judgment. Variks provided the Reef with Skolas' plan to assault the military fortress of Cybele, leading to the capture of Skolas and most of his lieutenants, bringing an effective end to the Reef Wars.
Meeting The Guardian
While tasking his Crows, Uldren ordered one to watch the entrance of the Black Garden and to follow any who passed through its gate. As the Crow left to set about its task, Uldren noticed one of the drones flying to avoid him and swiftly caught it in his grasp. Questioning it, he learned it was tasked by the Queen and forced it to state its task by reminding the Crow that it served him. It revealed that it was monitoring the Black Garden and that the Black Heart and the Sol Progeny which guarded it were growing stronger. After a brief moment of contemplation, Uldren brought the Crow with him to his sister's throne room and presented it to her, while noting that he believed he had earned the right to share this news.
When a Guardian came to Reef asking for an audience with Mara, Uldren greeted them with hostility and told them that he did not believe they should be granted access to the Queen. As the Queen's Fallen guards entered the throne room, the Guardian panicked and grabbed a gun from one of the Awoken guards, causing Uldren to draw his knife and hold it to the Guardian's throat. Mara then entered the room and explained that the Fallen were hers, and after the guard retrieved their weapon Uldren removed his knife from the Guardian's throat and apologized to the Queen for the disturbance. When the Guardian declared they were looking for the Black Garden, Uldren was surprised, but was unimpressed that they intended to destroy the Darkness at its heart, dismissing their goal as unimaginative. The Guardian asked for help entering the Garden, and Uldren questioned why they would help them. Mara then called her brother over to the throne, and suggested that the Guardian retrieve a Vex Gate Lord's eye so they could make a key for them. Uldren was pleased by this suggestion, as he believed the Guardian would perish attempting the task.
Upon the Guardian's return, Uldren believed that they had run from the Gate Lord and mockingly reassured them that their was no shame in their action. The Guardian was defiant and said they did not run, transmatting the head of a Gate Lord into the throne room. Uldren was stunned they had succeeded, and when they asked for the key and location of the Black Garden as agreed, he angrily noted their ignorance of its location to his sister. She told him to provide the key and coordinates, confusing and angering him further, and Uldren exclaimed that if she wanted the Guardian dead they should just kill them now. His sister informed Uldren that his guessing of her motives was only revealing his own, and he bowed before her and claimed he only wished to serve his people and her. After she directed him once more to gift the Guardian the eye, Uldren reluctantly pulled out his knife and pried the eye from the Gate Lord's head. As he did so, the eye lost power, and Uldren cheerily noted that all the Guardian's efforts were wasted. Mara disagreed and ordered him to hand over the eye anyway, and after a moments hesitation he tossed them eye and provided the coordinates of the Black Garden's entrance on Mars. Mara then proclaimed that the Guardian would come to the Awoken's aid if they should ever need it, which Uldren bluntly noted meant that they owed them for this. As the Guardian left, Uldren sarcastically wished them luck getting through the Cabal's exclusion zone on Mars.
The Wolf Rebellion
Several months after aiding the Guardian in their hunt for the Black Garden, the House of Wolves rose in rebellion against the Reef after Skolas was freed by the Nine and led a prison break from the Prison of Elders while also declaring himself Kell of Kells. Uldren and his Crows gathered intelligence on the Wolves' movements and kept Mara, Petra Venj, and Variks, the Loyal apprised of their actions while they worked with the Guardian to hunt down the rebellious Fallen. After the Guardian slew several members of the Silent Fang on Earth when they attempted to assassinate leaders of the House of Devils to let Skolas conquer them, Uldren met with his sister in her throne room. While she was pleased by the deaths of so many Fang members, Uldren informed her that he found nothing about the situation amusing, as his Crows had learned most of the Silent Fang was still alive and all had escaped from the Prison. His sister reassured him that she was not amused either, and ordered him to have Petra set more bounties on the Fang to ensure that none would continue to live.
Shortly after meeting with the Queen, Uldren was visited on the Hull of Crows by Petra, Paladin Yasmin Eld, and Techeun Shuro Chi to discuss Skolas and the Wolves' next actions. He informed them that a meeting had been set between Yavek, Wolf Baron and two Barons from the House of Kings, Vekis and Paskin. He explained that the Kings ruled the Devils from the shadows and had nearly conquered the Last City twice, and would thus never simply bow before Skolas' claim to be Kell of Kells, as the mysterious Kell of Kings had his eyes on the Traveler. Petra and Venj were skeptical, questioning Uldren on if Skolas managed to trick them with power or an artifact, but the Master of Crows believed nothing short of the Traveler proclaiming it could convince the Kings that Skolas was worthy and blunt their ambition. This statement cause Petra to note Uldren sounded like he admired the House of Kings, and he stated that any power cleverly used was worth admiring. Yasmin asked what the Baron's objective was then, and Uldren explained that he suspected they were in fact assassins sent to kill Yavek as a message from the Kell of Kings to Skolas. Shuro warned Uldren to ensure his assessment of the situation was free from personal bias, and Uldren demanded to know what she meant by that.
Uldren later reported to Mara that with all of Skolas' efforts to bring the House of Kings and Devils under his banner denied, he had returned to the Ishtar Sink on Venus. His sister guessed that he intended to rule from the deceased Winter Kell Draksis Ketch, the Simiks-Fel, but Uldren noted that while he had thought the same, his Crows reported that the Wolf Kell was not there. He then informed her that Skolas' Kell's Guard were spotted leading raids into the Vault of Glass in search of Vex technology. Intrigued by this development, Uldren was ordered by the Queen to have Petra bring Skolas back alive for questioning.
After Skolas was captured, Uldren travelled to the Prison of Elders with Mara and the Techeuns to view their prisoner. As they and Variks stood over the would be Kell of Kells cell, Uldren was amused by Skolas' attempts to reach them through the bars. However, Skolas began muttering something which Variks translated as "Dark-Binder" and "Light-Snuffer". This caused Mara to begin to leave, but Uldren questioned what her sentence for Skolas' crimes would be. She gave Skolas to Variks for the Guardians to fight in the Prison of Elders, and as Uldren held the door open for her to leave he was ordered to send Crows to Osiris on Mercury and Eris Morn at the Tower to tell them of what Skolas' had said.
Osiris and Eris soon arrived at the Reef to meet with Uldren and Mara to discuss the coming of Oryx, the Taken King. When Eris arrived at the throne room, Uldren noted that she did not have a Ghost, and she informed him that her next death would be her last. He dryly noted that his would be as well and took his place before the Queen's throne. As the meeting began, Uldren narrowed his eyes in disapproval at the way Eris spoke to his sister but did not speak out, but the Queen did not care about the lack of formality and instead declared that all of them would form a plan to kill the Taken King.
Search for the Queen
He participated in the Battle of Saturn and was the only known survivor at the time, as his ship crashed on Athabasca, Mars. However, his survival was unknown at the time, as control of the Crows passed to Variks; griefstricken Awoken even comissioned a sword in his honor. Uldren suspected that his sister survived the battle, as he could still sense her. A group of Crow drones found him and repaired his ship, but when he tried to break the atmosphere, he was shot down by Cabal and crashed in Hellas Basin. With all his Crows destroyed, his ship wrecked beyond repair, he treks across Mars looking for a way to get off-world. It is in these moments that the grief of losing to Oryx wears him down, causing his faith in her plans to fade at the same time his feel for her presence fades. However, when the sensation of her survival returns, he spends the rest of the Taken War and the SIVA Crisis searching for her, evading everyone including his own followers. His last known report indicated he allowed himself to be captured in order to find the Kell of Kings, to raise an army via some unclear means. He also indicated that his sister approved of his actions. Eventually, he is brought before the King's Kell, who submits to him and asks for his help in uniting the Fallen. Hoping to use their resources to search for Mara, he agrees.
The United House
Following Craask's plea, the former prince put out the call to rally the scattered Fallen Houses across the Solar System. In the year following the SIVA Crisis, Fallen everywhere began collectively retreating from all of their known strongholds to answer Uldren's call, leaving their banners behind as well.
The Mad Traitor
After the Red War, Uldren's alliance with the Kings lead him to become a traitor to his people. At some point, he learned that the Broken Legion was a trojan horse set up by the Red Legion. Rather than reach out to Petra, he allowed the Red Legion to invade, further destabilizing the Asteroid Belt. With the help of his Fallen allies, he began a campaign against what was left of his home in order to distract Petra from collaborating with the City. During the raids, he killed his people, stole from them, and brought more destruction to his home, using Mara's sacrifice at Saturn to justify his treachery. At the same time, between the strengthening of the Light brought about by the Traveler's awakening, and the fact that he could no longer hear Mara's voice, Uldren was driven to the brink of madness to the point where he consumed Ether to numb himself. On top of that, some unknown ailment was damaging his eyes, later made apparent to be a Taken infection, most likely as a result of the battle with Oryx.
On an outing, Uldren faintly detected Mara's presence, which led him to Fikrul, who was barely alive after being ambushed by Guardians. Uldren managed to save the Archon by unconsciously making a wish to Riven, causing Fikrul's Ether to become corrupted, causing Fikrul to see him as some sort of father-figure. As time goes on, Uldren found a kindred spirit in Fikrul, and they bonded through their shared dislike for their peoples' fates, as well as desire to see them evolve through extinction. He decided to team up with Fikrul and take over the Kings fully. With the help of the Scorned Barons, Uldren took over the House of Kings, and had his new right-hand dock the former Kell, reducing his rank to Dreg. After this, Uldren left the Scorn to their devices and resumed his search for Mara. He decided to try talking Petra over to his side, but after finding her in Thieves' Landing, he sadly saw that Petra has, in his eyes, become irredeemable. In that moment, "Mara" contacted him again. After the sighting, he tracked down Illyn and accused her of trapping Mara in The Dreaming City. Illyn denied it, having given up on Mara being alive, and accused Uldren of madness. While he was ranting about his plans for the Awoken's future, he had a break that moved him to surrender to his people for his crimes. Seeing that something was was wrong with him but unable to excuse his crimes, Petra, Cayde-6, and a strike team threw Uldren into a cell in the lowest parts of the Prison of Elders.
A year following the Red War, he and Fikrul took advantage of the Warden Variks' wavering loyalty and tricked him into releasing the prison's inmates, causing the Scorn to breakout out en-masse. With the united Scorned Barons, they faced down the Hunter Vanguard Cayde and destroyed his Ghost, Sundance, where Uldren then finished off with his own Ace of Spades, fleeing the scene afterward. From his perspective, he was doing something noble. Uldren's ultimate motive is to find his sister, Mara Sov.
With the aid of the Fallen Scorn, Uldren Sov sent the last remnants of the Reef into disarray and scattered the Awoken. He claimed to know that his sister still lived and communicated with him, telling him that she was imprisoned within the Awoken's secret structure, the Watchtower. To release her, Uldren required a combination of Light and Darkness to open the gateway for Mara Sov to be free. Already infected with the Darkness, Uldren sought out the Light in the European Dead Zone, and recovered a Shard of the Traveler but narrowly escaped an assassination attempt by Petra Venj.
Time was soon running out for Uldren, as each of the Scorn Barons fell to the Guardian one-by-one, leaving only the Fanatic as his last defense as the Guardian closed in. The Fanatic soon fell, but at the center of the Watchtower, Uldren opened the gateway with the Darkness within him and the Light Shard but instead of his sister appearing, a grotesque servitor-like Taken creature, the Voice of Riven, took her place, revealing that Mara Sov was never speaking to him but it was instead, Riven using him to free itself. Uldren realized this too late as the Voice of Riven swallowed him whole.
However, the Guardian soon fought their way to the center and fought with the Voice of Riven, and despite the creature's strange powers, the Guardian succeeded in destroying it, releasing Uldren. The Guardian, with the Ace of Spades, approached a defeated Uldren who questioned where his sister was, but Petra Venj soon arrived at the scene, equally ready to execute him. Uldren adamantly claimed that everything he has done was for Mara, but both the Guardian and Petra were not convinced. Ghost, however, stated that simply executing him wasn't right, but Petra angrily stated that she knew what Cayde-6 would do. Uldren further supported this question on what Cayde would do and asked the Guardian on what they plan to do. After a moment of silence and thought, they aimed the Ace of Spades at him. Uldren closed his eyes and was promptly executed, avenging Cayde-6 as well as the Awoken that Uldren had forsaken.
The Crow's Biography
Uldren was eventually laid to rest in the Dreaming City on a dais covered in a sheet as per Awoken custom, where he remained for a short amount of time. Soon after his death, a Ghost named Pulled Pork found and resurrected him as a Lightbearer. Uldren sat up in shock as the Light returned him to life, although he was unaware of who and where he was.
A little less than a year after his revival, sometime during the Dawning, Uldren was seen camping out in a shipping container with Pulled Pork. He has yet to go to the City and spends his days alone, save for his Ghost. All the Guardians he's ever encountered have shown a range of negative responses to seeing him, from grief to outright hostility.
Caught in the Spider's Web
Due to Crow's naivety he ended up forcefully working for The Spider by him promising Crow safety from the outside world. However soon Crow would be forced to work for Spider due to Glint being lined with explosives which could be detonated at any time if Spider felt he was going rogue. From there Crow was used for constant menial tasks that Spider would normally pay his Associates to do. Crow was originally oblivious to Spider's exploits however the longer he observed his Associates, House of Dusk Fallen, and even other Guardians, he began to doubt Spider's want to "Help" Crow and Glint, eventually wishing to flee from Spider, but couldn't due to Glint's explosive shell.
The High Celebrant Rises
Once Season of the Hunt begins, Crow is tasked with being the researcher of Wrathborn, using technology within Spider's safehouse to jury-rig lures to give to Guardians in order to lure out and hunt Wrathborn leadership. While the hunt continues, Crow manages to augment the lure in a way to hunt down the leader of the Wrathborn, Xivu Arath's High Celebrant. He then provides The Guardian with the special lure to kill the Celebrant.
Once the hunt begins Crow heads to the Dreaming City's Ascendant Plane in order to corner the Celebrant and force them back to the Dreaming City so the Guardian can continue the hunt. During the hunt Crow is trapped within the Ascendant Plane and crippled, forcing the Guardian to head in and finish the celebrant themselves. While the Guardian struggles to keep the pressure on the Celebrant, Crow recovers and finds a way to rig the Celebrant's portals, forcing them to stay in the Ascendant Plane and stun them, allowing an opening for his partner to end the Celebrant's campaign and its life for good.
Dreams of Light
As the hunt went on, Crow began experiencing strange, vivid dreams, and confessed in Glint about it. Soon enough the Guardian found several paracausal feathers in places Crow had dreamt about, and discovered a golden hawk which led them to a previously unknown shard of the Traveler in the Hallowed Grove. They fought off the Taken forces of Savathûn, the Witch Queen, and the Traveler gifted them with a reforged hand cannon the Hawkmoon. They celebrated the victory over a bonfire, with Savathûn apparently watching them.
The hawk would call them to the shard a few more times, and during one such instance, the Guardian stopped by Crow's nest in the EDZ and discovered his old ship had been replaced with a new one. The Traveler would gift a ship to the Guardian too: Radiant Accipiter, paracausal in nature, much to Amanda Holliday's bewilderment.
Slaying the High Celebrant
After weeks of hunting the Wraithborn and picking up pieces of the High Celebrant's trail, Crow informed the Young Wolf that he had tracked their prey to the Dreaming City. Providing the last upgrade needed to attract the High Celebrant's attention with the lure, Crow smiled and stated that they were, as the Spider liked to say, in business and ready for the hunt. Traveling to the Dreaming City with the Young Wolf while Osiris provided overwatch, Crow found the High Celebrant and followed it through a portal into the Ascendant realm while the Young Wolf got into position with the lure before a cryptolith. Crow noted that he had been studying how the Celebrant moved between planes and was learning its tricks. As the lure was placed Crow attacked the High Celebrant and wounded it so that it bled energy to track easier as it traveled between realms. After the Young Wolf forced it back into the Ascendant realm their Ghost reported it left residual energy behind. Crow explained that they could use that to open a gateway and continue their pursuit of it. He remained in the Ascendant realm while they pursued the High Celebrant back to the Dreaming City, stating that as long as they were within the same spot in both realities the High Celebrant would have nowhere to hide. He then drove several Fallen Wraithborn into Dreaming City so that the Young Wolf could slay them and use the reclaimed Light energy to hurt the High Celebrant.
However, the High Celebrant soon realized what they were doing and healed itself, no longer bleeding energy for them to track. It fled back to the Ascendant realm and sealed the way back, and without the energy it left behind Crow and Glint were left with no way to return. With the tables now turned, Crow was pursued by the High Celebrant through the Ascendant realm. It caught up with him and after a short fight broke both his legs and flung him into an abyss. He sent a message to the Young Wolf asking them not to blame themselves if things went badly for him and thanking them for giving him the choice the help and feel like he was doing some good while trapped in the Spider's employ. The Young Wolf and their Ghost managed to recreate the portal into the Ascendant realm and messaged them, although Crow stated he could handle the situation. Glint disagreed and revealed his injuries, although he quickly healed Crow. Crow insisted he was fine and had a plan, directing the Young Wolf to the High Celebrant's location.
Returning to where the Young Wolf had entered, Crow asked them to keep it busy while he returned to the Dreaming City. Ghost questioned if he was leaving, but Crow noted that the High Celebrant had to die in the Ascendant realm to truly be defeated and he believed that the Young Wolf had the best shot at making that happen. Waiting at the point where he had deduced the High Celebrant would emerge when it retreated back to reality, Crow used what he had learned of traveling between realms to close the portal shut on the Hive warrior, sending it back into the Ascendant realm and vulnerable to being finished off. Osiris was astounded as Crow explained what he had done and asked the Young Wolf to recover the High Celebrant's head to present to the Spider as proof their mission was a success and told them it had been an honor working with them.
The Bird Flies Free
Returning to the Spider's lair, Crow and the Young Wolf reported their success to the crime boss. Spider told the Young Wolf to name their reward, having promised anything within his lair, and Crow was stunned when the Young Wolf named his freedom as their prize. Spider balked as his guards readied themselves for a fight, but the Young Wolf reminded them of their deal as Crow nervously watched. Trapped by his own wording, Spider agreed and told Crow to fly away before demanding they leave his lair. Exiting the building, Crow and Glint wondered what they would do now and that their freedom did not feel real. Turning to the Young Wolf, Crow questioned why they had done this for them, and their friend told them that it was because the Crow was a Guardian.
Crow returned to the lair to have the Spider remove the explosives from Glint and resolved to remain around of his own free will to finish off the remaining Wrathborn. When the Young Wolf visited him Crow wearily noted that he was suspicious of the Spider holding up his end of the bargain and noted he never did anything if he did not get something out of it. He informed them of his plans to remain for a few weeks to wrap up his business and that afterwards he would find some way to enjoy his freedom. Several weeks later, Crow was visited by Osiris and inquired about his plans. Crow stated he was planning on exploring remoted places across the Sol System, including Venus, but the former Warlock made him another offer: to come the Last City and join his fellow Guardians in the Tower. Agreeing to the plan, Crow began packing his belongings and making ready to depart from the Shore. Before leaving, the Young Wolf visited him one last time in the lair. He explained to them his plans and expressed nervousness about leaving the Shore and going to the Tower. However, Crow resolved that while he might be judged for who had been in his previous life he would not cower in fear of it and would act like a Guardian.
Travelling to the European Dead Zone on Earth to rendezvouses with Osiris, Crow pondered what life was going to be like in the Last City. He wished he had the bottle of alcohol that Druis had once offered him as he had never been drunk. Glint suggested he could drink as much as he wanted in the City, but then suggested caution about that after quickly looking up details on what happened when drinking. Crow told Glint he suggested caution about everything, but became nervous as he thought about all the people in the City who could recognize him for crimes he did not even remember. Glint reassured him that he could do whatever he wanted as long as he remained in disguise. Crow expressed a desire to eat food that did not taste like ether and gun oil and was startled when Osiris arrived and suggested the Long-Boy Special at the Drunken Noodle. Remarking that he had seen images of noodles and that they reminded him of worms, Crow listened as Osiris suggested they get him a change of clothes before going to the City. Glancing at his cowl emblazoned with the Spider's symbol, Crow agreed and followed Osiris back to his ship where a change of clothing awaited him. As they boarded, Osiris halted and warned that a day would come in which Crow's face would be seen and the City would know who he was, but reassured him that one exile to another that he could trust Osiris when that happened. Happy with Osiris's support, Crow began to feel like he might find acceptance after all.
Clash of the Chosen
After arriving in the Last City, Crow was gifted a new suit of armor that was paid for by Osiris, although Glint chose the material it was made from. He also took to wearing a mask to hide his face for his own safety. He was asked by Osiris to join a task force put together by Commander Zavala as a reconnaissance expert. The task force was dedicated to preventing Empress Caiatl of the Cabal from reuniting the Cabal Empire with the remnants of the Red Legion. Crow agreed to join and took up residence in the H.E.L.M., where the task force was running its operations from.
Contacting his assets in the Tangled Shore, Crow learned that most remaining Red Legion commanders were brutes and warlords looking for ways to impress Caiatl. Crow met with Osiris and the Young Wolf to discuss the situation. While Osiris cautioned him to ensure he kept his mask on at all time, Crow reassured him that would not be an issue and that if it was the price he had to pay for making a difference for humanity he was fine with it. Osiris reassured him that he once had to walk in disguise within the Last City and that Crow would be welcomed like he was one day. Turing their attention to the Cabal, Crow commented that Saladin would hold the frontline as they identified and eliminated targets of importance. Relaying what he learned on the Shore, they made a plan to eliminate all contenders for Caiatl's War Council and prevent the Cabal from reorganizing for war. With their plan in place, Osiris dismissed them, but before departing Crow commented to the Young Wolf that they would be working together in the field again and that he was getting the impression they liked having him around. Crow began his part in the operation by spying upon a Cabal operation on Europa led by Basilius the Golem, learning that he planned on gathering Golden Age technology to augment Cabal soldiers as an offering to Caiatl. Crow reported his findings to Saladin, who made Basilius the first target of the task force.
Crow decided to visit one of the Spider's betting outposts in the Cosmodrome to gather more information on the Cabal's activities. As they approached, Glint worried that it could be a trap, but Crow reassured him that if the Spider wanted him back he would have made his move already and that the outpost had no idea he was coming. Several snipers suddenly trained on him, and Crow quickly shouted the password. The snipers stood down as Avrok opened up a nearby hidden door. Entering, Crow noted that the associate had been promoted and questioned what the odds were on Caiatl winning. The warehouse was dark, and Crow's eyes had trouble adjusting to see what was stored within. Avrok growled that information cost, causing Crow to produce a bundle of glimmer that partially lit up the room and allowed him to see numerous Cabal weapons, vehicles, and technology being used as collateral by betters. Commenting on the objects, Crow's attention was caught by one Sparrow, which Avrok noted belonged to Val Ma'rag, who had bet he would become the first member of the War Council. With this new information, Crow turned back to Avrok and placed his own Sparrow on a counter-bet, claiming Ma'rag could have it if he ascended to the Council, but suggested to Avrok that the Guardians would disrupt the Val's Rite of Proving. Avrok was pleased with the bet and to do further business with Crow, departing to get the betting book. Glint was concerned about losing their Sparrow and noted Crow hated walking, but Crow simply shrugged and stated he liked winning more.
As the operations against the Cabal continued, Crow developed a friendship with Amanda Holliday, a Shipwright providing aerial support to the task force. After one operation on Nessus, Crow wondered why the Cabal had deployed a tank on Nessus and Amanda claimed all they knew was brute force. He questioned where she had learned Cabal tactics and learned from her that she had fought on the frontlines of the Red War and witnessed the Cabal slaughter defenseless civilians and Lightless Guardians. Crow was surprised, having little knowledge of that conflict, and did not understand why the Cabal still fought even after losing their leader Dominus Ghaul to the Traveler.
The Psion Conspiracy
Crow subsequently travelled to Nessus and learned that Psion Ixel, the Far-Reaching was leading a group of Cabal on incursions into Vex territory seeking their prediction engines to offer knowledge of the future to Caiatl. Although Ixel was killed, three other Psions escaped with data stolen from the Vex prediction engine. Crow was dispatched to monitor Cabal reaction to their strikes and observed that while they were angry with the Guardians disrupting the Rite of Proving, many others were upset that Caiatl had tried to recruit humanity into the Cabal to begin with. Returning to the H.E.L.M. to meet with Osiris and the Young Wolf, Crow reported his findings. While Osiris saw this as an angle to be exploited, Crow voiced concern about retaliation by the Cabal using the data stolen from the Vex, which they had determined revolved around the future of Commander Zavala. Osiris noted that it was only possibility not fact they found and assigned Crow to work with the Young Wolf to stop whatever the Psions were plotting. Crow protested not informing Zavala of what they found, but Osiris expressed hope the two of them would ensure there was nothing to report. Turning to the Young Wolf, Crow stated that he would take that as a compliment to their abilities and departed to begin his new mission.
As Crow began investigating behind the Imperial Cabal's lines for signs of the Psions, Saladin assigned the Young Wolf to disrupt more Rites of Proving to distract from Crow's operations. With the aid of the taskforce, Crow was able to pick through the information he gathered and the data recovered by the Young Wolf and realized that all of the futures that the Psions had looked at involved Zavala's demise. Believing that the Psions intended to assassinate Zavala on behalf of Caiatl so that the Empress could coerce the Guardians into her service, Crow summoned the Young Wolf back to the H.E.L.M. and shared his concerns with his friend. Noting that Glint needed more data to accurately pinpoint when the assassination would take place, Crow requested that they gather their own data from the Vex's systems. The Young Wolf chased down several leads gathered from the Vex data and recovered a mysterious device from a Cabal encampment on Earth. Crow met with them once more and informed them that the device would disrupt the Light of a Ghost it was attached to and render their Guardian vulnerable to a final death, a refinement of the cage used by the Red Legion on the Traveler. He noted the device was an elegant way of defeating Guardians without a fleet or army, but a single assassin. Crow passed the information along to Osiris and vowed to keep an eye on Zavala to ensure his safety. He was pleased that his hunch was right, but quickly recalled that several Guardians had been reported missing over the past few days, and the body of one had been found with no Ghost or Light and was seemingly killed by human weapons. This caused him to suspect that these Psions were experienced and had been planning this for some time.
For the next week, Crow and Glint discreetly watched over Zavala. During the day he covered him from after with a sniper rifle, while at night they snuck into the Tower and served as secret bodyguards due to the more limited visibility. As they watched over the Commander one day, Glint suggested he should get a new mysterious shell from Tess Everis since they were undercover, but Crow pointed out that they were on a reconnaissance mission and that only Crow needed to hide his identity as no one knew who Glint was. His Ghost protested that he knew everyone and had been around for hundreds of years. Crow teased him that they knew him as "Pork Bun" and were not aware he had a new Guardian. As Glint grumbled, Crow adjusted his position deeper into the shadows to ensure that Zavala did not spot any reflection from his binoculars, closing his cloak deeper around him. As he admired the garment, he felt guilty for being stringy after the generosity shown to him, and told Glint they would get him a disguise shell after the mission. Glint was ecstatic while Crow joked that it would only be a rare or legendary shell and that he would need to find another Guardian if he wanted an exotic one. Glint refuted that notion and told him he was the best despite what Saladin said, causing Crow to snort and note that he hoped the Iron Lord would realize they were all on the same side soon and treat him like a real Guardian.
One night while watching over Zavala in the Last City's first garden, Crow removed his mask for a moment to relax. However, he spotted a Psion approaching the distracted Commander and shouted a warning, enabling Zavala to spin around and kill the assassin before they could strike and allowing Crow to quickly depart. Crow met with Osiris and the Young Wolf in the H.E.L.M. to discuss the thwarted assassination, where he learned that Zavala had seen enough of him to identify his face but was passing it off as a hallucination. The two argued over Crow's actions, and he insisted to Osiris that the situation was far more deadly than the Warlock realized and that he had to risk his cover due to Zavala being distracted and his Ghost Targe being out and vulnerable as the Psion approached. Osiris criticized him for being unmasked when he did so and questioned if the Spider had not taught him that small mistakes could have large consequences. Crow angrily shot back that he supposed Osiris had learned that chasing Xivu Arath. After a moment of silence Osiris warned him to choose his next words wisely. Calming down, Crow acknowledged that he should have kept his mask on but did not regret acting and that he still believed Zavala was in danger. Osiris agreed and ordered Crow to become Zavala's new bodyguard, noting a full security detail would be too obvious and they needed to draw in the assassins, but warned him to never speak and always keep his mask on. Crow reassured him that he would handle the assignment with the utmost discretion.
Before departing, Crow spoke with the Young Wolf and explained what had happened, regretting that he let his guard down to feel safe for a moment. He knew that whatever choice he made in that moment, to either let Zavala die or expose himself, would invite criticism from someone but did not regret his choice. Ruminating on how he was sure the Young Wolf had lost people in similar situations where they were too far to act but close enough to wonder, Crow bet that they would given anything to stop it no matter the cost. Recalling his own experiences at being at the mercy of other Lightbearers attacking without knowing the reason why or how to stop it, Crow declared he had no intention of ever being the cause for more pain. He vowed that the next time an assassin came for Zavala they would have to go through him, mask or not, before quickly requesting that they not tell Osiris he said that last part.
A Growing Friendship
Following another operation on Nessus, Crow noted Amanda was tense and more aggressive towards the Cabal than normal due to the failed assassination attempt on Zavala. He tried to reassure her that he understood how she felt, but noted that the Commander had survived. Amanda retorted that Zavala was all that was holding the City and Vanguard together and that if the Cabal had killed him, then they would have been doomed. Surprised by the passion and hurt in her voice, Crow questioned how close the two were, and Amanda explained that Zavala and Ikora were like family to her, just as Cayde-6, the deceased Hunter Vanguard, had been. Crow recalled the stories he had heard of Cayde and wished he could have met him, causing Amanda to state that at least his killer got what was coming to them. Expressing sympathy for her loss, Crow was caught off-guard when she asked him to get a drink together when they returned to the City. He told her that he had always wanted to do that, surprising her and causing Amanda to warn him to get ready for a fun time.
As they returned to the Tower from another mission, Crow inquired about a tune Amanda sang as she landed their jumpship, which she explained came from her father during their time on the road. As they exited the jumpship, Crow noticed Osiris having an intense conversation with a Exo Titan and quickly pulled Amanda back behind a landing strut. They both winced as Osiris yelled at the Exo to never mention "her" again. Looking to Amanda for an explanation about what they witnessed, she promised to tell him later but went over to comfort the Titan. Crow worriedly followed, as being introduced to new people still made him nervous even with his mask. Amanda introduced him to Saint-14 and told Crow he ran the Trials of Osiris, surprising the Hunter, as he did not think the exuberant Titan matched what he had heard of the Trials. As they talked, Crow wondered what a dour figure like Osiris could possibly have in common with someone like Saint. Saint departed for a meeting with Ikora, leaving Crow with barely enough time to bid farewell, much to Amanda's amusement. He questioned if she could explain now, but she would only do so over a round of drinks, insisting that Crow buy this time.
Crow accompanied Zavala to the sealing of an armistice between the Vanguard and the Cabal Empire on Nessus after the Young Wolf led a strike team in defeating Caiatl's chosen champion Ignovun in a final Rite of Choosing. As the ceremony progressed, the participants were caught off-guard when Zavala's Ghost Targe was suddenly imprisoned in a Light-draining cage. As Zavala collapsed, disoriented and Lightless, the Psion attendant Valir charged the Commander with the ceremonial blade being used to ritually seal the armistice. Crow moved to stop the Psion but failed to stab him, only intercepting the first blow and fell to the ground injured and maskless. However, Caiatl personally intervened and killed Valir before sending off one of her bodyguards to find the sniper who imprisoned Targe. Crow reached out to Targe and broke the cage off of him, restoring the Light to Zavala. The Commander inquired if he was alright and Crow stated that he was alive, but reluctantly turned to face Zavala without his mask. Although Zavala was shocked to see his face, Crow was equally surprised when he reached out and offered him a hand up, which Crow happily took.
Returning to the Last City, Crow and Zavala discussed his situation, with the Vanguard opting to keep his identity a closely guarded secret that they would spread slowly while Crow continued to lay low. However, Crow's actions also made him a hero of the Tower and earned him an offer of official institution as a Guardian and recon agent for the Vanguard. For his first assignment from Zavala, Crow was asked to hunt down the other Psion conspirators behind the treaty incident. Now maskless, Crow met with the Young Wolf at the H.E.L.M. and informed them of the good news. He believed Caiatl's word that she was not behind the assassination attempts but was not willing to take chances on more happening. Reflecting on his interaction with Zavala, Crow noted that the Commander saw him, not who he was before. Noting that while it was taboo to discuss who they were before, Crow now felt that the weight he had around his neck since being reborn was finally lighter. Bidding his friend farewell for now, Crow was sure that they would see each other again soon.
Personality and Traits
Prince Uldren Sov has proven that while intelligent, cunning, clever and loyal to his sister and the Reef, he has shown to be condescending, smug and arrogant towards outsiders and enemies, particularly to the Guardians. Even some members of the Reef dislike him for his negative attitude but can't deny his skills, leadership, and expertise. His leadership of the Crows in both the Reef Wars and the Wolf Rebellion showcased his skills in intelligence gathering. In addition, he proved resourceful enough that he was able to avoid the detection of his own followers. During the chaos within the Reef, Uldren did show further, albeit ruthless, leadership skills as the Fallen Scorn followed his orders without question. Even the Barons held a sense of reverence for him.
Uldren has always disliked all Guardians, primarily because of their complete disregard of their place in the grand scheme of things. To that end, Uldren makes it his mission to antagonize nearly any Guardian he encounters. For his own amusement, he also habitually states that "Rasputin shot the Traveler" during The Collapse just to rile them up. When The Guardian first came to the Reef, Uldren largely didn't want to help the Guardian in finding the Black Garden despite the threat of the Vex and was content to have nothing to do with them. He also doesn't think much of Guardians' abilities, as he quickly suspected that the Guardians would fail in the mission to find and defeat a Vex Gate Lord. When the Guardian actually succeeded, he was left shocked and almost livid that he was proven wrong. In addition, Uldren is confused and angered with his sister's growing alliance with the Guardians. Out of loyalty, he kept his opinions to himself and followed orders obediently.
What happened after the Battle of Saturn with Oryx and surviving is unknown but Uldren has become even more ruthless and cruel. He was willing to betray the Reef, ally with the House of Kings and later the Fallen Scorn and commit murder in cold-blood to meet his goal of meeting his sister again. Unfortunately, Uldren's desperation to find his sister made him susceptible to Riven's machinations and his corruption of the Darkness after the Battle of Saturn only made it worse for him. He began to see visions of Mara Sov, directing his actions, which made him act erratically to those who knew him well. His condition only grew worse when Riven's hold over him strengthened, causing him to willingly betray and kill his own people. However, while his actions and mannerisms remain relatively the same to the Guardian, Uldren showed clear cases of conflict within him, especially when the Guardian traveled through the Ascendant Plane within the Watchtower. While he longed to be reunited with his sister, part of him believed that something was wrong. However, he found this out too late and was horrified when he realized that his sister was never the one speaking to him.
After being freed and defeated after the Voice of Riven's demise, Uldren remained self-righteous in his goals, adamantly stating that all of his actions were for his sister. Ultimately, when his own death came for him, he nonetheless accepted his fate.
Following his death and resurrection as a Lightbearer, Uldren's memories had been wiped clean. He has shown to be nothing like his past self, becoming a more soft-spoken and kindly figure. Before joining the City, any Guardian he encountered in the field would react to him with hostility or grief, and although he doesn’t know why, he does understand that his past life must committed grave atrocities to earn their ire and is adverse to finding the full truth. It is because of this that he also strives to help out wherever he can to prove himself as a good person. He has shown to look up to and admire The Guardian. He also shows sympathy for the plight of some of the enemies of the last city, particularly the Fallen and the Cabal. During Season of the Chosen, he displays a more snarky side to him, particularly during some of his interactions with Osiris and Saladin Forge, in the latter case even sarcastically calling him out on his absence during the Red War.
The H.E.L.M., The Last City, Earth (Season of the Chosen) 
Season of the Splicer
List of appearances