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Following the arrival of the Black Fleet and the sudden disappearance of the Witch Queen, Xivu Arath sought to establish a foothold in the Sol System in her sister's absence with the aid of her High Celebrant. Acting as her envoy, the Celebrant summoned Hive structures known as Cryptoliths from the Ascendant Realm across the system to deploy an army of Wrathborn from across dimensions and corrupt a new army from the forces of the Fallen, Cabal, and renegade Hive.
They were majorly rampant across the Tangled Shore and the Dreaming City in tandem with the events on Europa. Their threat was challenged by Guardians working with Crow, "Osiris" and the Spider to drawing them out of the Cryptoliths and hunt them down. The High Celebrant was ultimately caught and eliminated in a final hunt by the Guardians and the Crow, driving back the Hive God's advance upon the Sol System for a brief period.
Later, they would be deployed to the Dreaming City by Xivu Arath who was seeking retaliation against her sister, who was held captive by the Awoken. Large numbers of Wrathborn along with forces of Taken and Scorn would form a collective assault on the City through the Ley Lines and the Ascendant Realm, seeking to capture Savathûn.
Wrathborn are mindless, enraged versions of their former selves, attacking former allies as readily as enemies. They are marked by a sickly green-glowing miasma that hovers around their heads and leave behind a trail of noxious soulfire that can be used to track them. They are given increased strength and resilience by the Hive magic of the Cryptoliths, making them an obedient army of monstrosities in service of the Hive god herself. They are also able to traverse across dimensions utilizing the Cryptoliths as conduits into the Ascendant Realm and must be drawn out with a Lure built by The Crow.
According to Osiris, who unbeknownst to everyone was Savathûn in disguise, the Wrathborn are perverted by Xivu Arath's magic beyond recognition that they are neither considered Hive, Fallen nor Cabal. One of the first Wrathborn Cabal was shown to have mutated abnormally given time; with its Severus cleavers fused to its hands and bladed tendril mutations growing out of its back and stomach. They also aid in the creation of more Cryptoliths by planting their own grown-tendrils into a shrine of black twisted spines, which are kept grown with the bodies of victims impaled upon their apex of crowns. Under Xivu Arath's influence, the Wrathborn are mentally stripped of everything beyond an innate desire to kill before their bodies are remade by the Cryptoliths into the monstrosities they are now.
Additionally, Savathûn (while posing as Osiris) also made note of the roles Wrathborn Fallen and Cabal had. Fallen Wrathborn, with their mobility, serve greatly for reconnaissance. The size and strength of the Cabal made them prime candidates for Throne Guards or beasts of conquest for Knights to feed upon. Psions corrupted by the Cryptolith would be brought into Wizard courts to serve as psychic lodestones for ceremonial dives into the Deep.
List of appearances