Omnigul, Will of Crota
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Omnigul, Will of Crota is an Ascendant Hive Wizard and the consort of the Hive god Crota; she is also the matron of the Spawn of Crota and the leader of his armies. She acts as the secondary antagonist for The Dark Below campaign, where she led the Hive in resurrecting Crota back to the material realm and raising an army to invade the City; both efforts were thwarted as she finally met her end by a fireteam of Guardians.
As the consort of the Hope-Eater, Omnigul served as the brood mother of Crota's spawn through their progenation, acting as keeper of the brood's worms and his chosen Will. Through their conception, Omnigul is also the mother of four daughters of Crota's direct lineage: Hashladûn, Besurith, Kinox, and Voshyr.
At one point she was the target of a then-unnamed Lokaar to replenish her strength using her death as tribute; which instantly invokes Crota's wrath only for Lokaar to be named by his father and raised to his court.
The First Crota Team
During Eriana-3's failed raid upon the Hellmouth, Omnigul was present when she encountered Eriana and the Hunter Sai Mota at the World's Grave. Their Light siphoned and losing her knives to the Hand of Crota, Sai engaged the Hive matron armed with only the bones of fallen Acolytes. Omnigul tore the Hunter apart forty-four times until only wormrot was left on her corpse to devour what remained of the latter's Light and Ghost.
The Dark Below
Several months after the destruction of the Black Heart, Omnigul was first sighted on Earth leading a contingent of Hive against the Fallen House of Devils before she met the Guardian during their hunt for Sardon, Fist of Crota, fleeing the scene as they wiped both parties. Eris Morn confirms to Ikora Rey her departure from the Hellmouth for the first time in years; with word having reached Eris about an imminent plot to sabotage the Warmind Rasputin and his Warsat defenses at the Seraphim Vault within the Cosmodrome. 
During the Hive's siege of Rasputin's bunker, Eris sends the Guardian to stop their advance from either destroying or corrupting the Warmind. When they reach the center of the bunker, however, Omnigul and her Watchers of Crota ambushed them by summoning waves of Hive reinforcements directly at the Guardian. When proven ineffective, Omnigul summons the Might of Crota before escaping yet again. Whilst it seemed that the assault to destroy Rasputin has failed, Eris believed that Omnigul has found what she sought from the Warmind.
Ultimately, Omnigul's efforts at bringing back Crota would be for naught as her summoning ritual was disrupted and her spawn hunted by Guardians, until she herself was left pursued by a Strike fireteam in an attempt to raise a new army beneath Skywatch. Despite her efforts at stirring the House of Kings against the Guardians, she was finally cornered at the Jovian Complex and killed, putting an end to her schemes. 
Shortly after the SIVA Crisis, Omnigul would make a brief return to Skywatch after having "undone" her death to continue her work, only to be swiftly killed again by the Guardian and their fireteam. Since Omnigul was capable of evading permanent death, Ikora remarked that the Vanguard will likely continue to face Omnigul as many times as it takes.
Following the discovery of the Pyramid beneath Luna's surface, the Hive under Hashladûn, a daughter of Omnigul and Crota, use its strange energies to create phantasm-like Nightmares. Constructing the eerie Scarlet Keep fortress near the Pyramid, Hashladûn aimed to use the Pyramid's power to exact her revenge against the Guardians for destroying her lineage. However, like her mother and father before her, Hashladûn would be defeated by the Guardians at the Scarlet Keep's highest tower.
Despite Hashladûn's death, Nightmares continued to appear across the system, with the stronger ones forming into some of the Guardians deadliest foes, Omnigul being among them. The newly formed Nightmare of Omnigul, Will of Crota, representing the Essence of Anguish, would become a target in the Guardians' Nightmare Hunts to contain the Pyramid's influences.
Omnigul is the only Wizard with an Arc Absorption Shield instead of a Solar shield. She can summon Hive and casts poison with greater frequency. Unlike normal Wizards, Omnigul fires her Darkness Blasts faster and more accurately than normal. However, unlike any other Ultra class enemies, she can be blinded by flashbang grenades, Ward of Dawn with the Helm of Saint-14 equipped, and the Purifier Robes.
Omnigul makes a return as the final boss of the Revisited version of the strike. Just like last time, she will stay in the center of the arena and suppress Guardians with her rapid fire Darkness Blasts as well as cast poison clouds for area denial. However, she will have locked the door that Guardians would normally run through for cover, trapping the players inside for the duration of the fight. The room will eventually get filled with a toxic gas that drastically slows the movement of any player that steps in it. Whenever reinforcements arrive, she will teleport away from the arena until all the reinforcements are dead. This will repeat until she herself is killed for good.
List of appearances