From Destinypedia, the Destiny wiki
Wyverns are bipedal Vex units, with double-jointed legs ending in clawed feet. Their most notable feature is a pair of fan-like appendages wreathed in glowing blue tentacles similar to Harpies, which project a shield on either side.
Wyverns make their first appearance on Europa and can be found in various activities such Strikes, Empire Hunts, Missions, Patrols, Lost Sectors, and the Spire Integration Public Event. Shielded variants appear on Nessus in Battlegrounds. They also make a non-cannon appearance in the Vault of Glass as Sacrificial Wyverns and feature prominently in Override activities.
They appear to be a hybrid of a Minotaur and a Harpy and have two forms. They battle in their bipedal form and occasionally switch to their aerial Harpy-like form to hover in the air and hurtle downward an aerial slam. The force of this attack can push all nearby non-Vex away and deals Arc damage.
They are highly durable and equipped with two shotgun-like Warp Lance turrets mounted on their bodies. This fires cross-shaped Void blasts that split into five smaller projectiles. They are also unique for fan-shaped appendages on either side, which if shot will create a shield to protect any Vex units behind it.
Wyverns also possess a Radiolaria core which can be tricky to expose, but achievable. By shooting its purple Warp Lance turrets near its eyes, Wyverns are briefly staggered to expose their core. This then grants players the opportunity to deal critical damage against Wyverns. Stasis elemental damage can also expose their core.
List of appearances