Wolf Rebellion

The Wolf Rebellion was a conflict between the Guardians of the Last City, their news allies the Awoken in the Reef and the House of Wolves who broke free from the Queen. Originally the Wolves were vassals to Mara Sov when she captured their Kell, Skolas, imprisoned their nobility and forced the remainder of their numbers into swearing allegiance in the aftermath of the Reef Wars. However, after the Queen sent Skolas to the Nine as a gift to placate them for sending the Crows into their territory only for one of their agents to release him and return him to lead his House. Leading a rebellion in the Reef to free his House, Skolas revealed he had far greater ambitions than just revenge on the Reef that imprisoned him but to make himself the ruler over all Fallen Houses and claim the Traveler from the Last City. In response, Mara Sov opened her realm to the Guardians to aid her hunting the Wolves.

Prelude
The prelude to the Wolf Rebellion began during the Battle of the Twilight Gap. All Fallen Houses banded together to seize control of the Traveler for themselves. The Devils, Winter and Kings houses met the Guardians in battle at the Twilight gap and the House of Wolves originally intended to join them and they could've turned the tide to the Fallen's favor but Queen Mara Sov intervened. Whether to save potential allies or for some other unknown reason, Mara Sov launched a surprise attack on the Wolves that claimed the lives of half their numbers and even their Kell, Virixas. Soon began a long and bloody conflict, the Reef Wars.

During a brief civil war between themselves, Skolas emerged as the new Kell and continued to wage war against the Awoken but his brutal tactics drove Variks, the Loyal to betray him at Cybele that led to his capture. With their Kell and remaining nobility captured, Variks named Mara Sov the new Kell of the House of Wolves. She gave the Wolves the choice of either submitting to her rule and live in the Reef or be destroyed. The majority of the Wolves chose the former.

Betrayal
During the Queen's search for Kaliks Prime, seeing if the Prime Servitor is still alive, she sent her spies the Crows out into the Jovians, angering her allies the Nine. To appease them, she sent Skolas to them as a gift but one of their agents freed him from his cyro cell and gave him a ship to return him to his house.

Gathering his followers from the Prison of Elders and leading an attack against the Reef, Skolas nearly assassinated the Queen before taking his house away to plan their next move. Angered by the Wolves betrayal, the Queen called in her favor to the Guardians, offering the riches of her realm for hunting down the Wolves.

"Prophesied Savior"
With Petra Venj and Variks as their guide, the Guardians tracked Skolas to Venus where he sought to gain the allegiance of the leaderless House of Winter. The Guardian was able to find Skolas rallying the Winter Fallen after calling himself the prophesied Kell of Kells, a Fallen warrior that would unite all Fallen Houses and claim the Great Machine. Though the Guardians were unable to capture Skolas and he was able to gain a number of the winters allegiance.

Traveling to Earth, Skolas sought to gain the allegiance of the House of Devils and House of Kings. While he set loose the Silent Fang to kill off the Devils remaining leaders, he sent one of his Barons, Yavek to negotiate with the Kell of Kings. However, the Guardians intervened as they killed a large number of Silent Fang assassins and killed Yavek and two King Barons, Paskin and Vekis.

Last Stand
Furious at the Reef and Last City interventions, Skolas desperately sought a means to defeat his enemies and force the other houses into recognizing him as the prophesied Kell. Skolas then had his splicers breach the Vex Vault of Glass to gain the weapons he desires. Fortunately, the Guardians once again intervened before they could gain any significant Vex technology, such as the Oracles.

With his enemies cornering him, Skolas sets up a final stand at the top of the Vex Citadel. The Guardians battle against the mad Kell who began to use Vex Warp Gates into bringing his house from across time to defeat the Guardians. Despite his efforts, the Guardians defeated Skolas into submission as they received orders from the Queen to capture Skolas alive. The Queen then sent Skolas back to the Prison of Elders, where he would soon meet his end, effectively bringing an end to the Wolf Rebellion.

Aftermath
For their contributions in the Rebellion, the Guardians earned their rewards, strengthened Reef-City relations and the House of Wolves were left leaderless, homeless and broken. The remainder of the Wolves then traveled to Mars to eke out a living, despite the heavy Cabal presence.

House of Wolves

 * Skolas, Wolf Kell
 * Beltrik, the Veiled
 * Drevis, Wolf Baroness
 * Silent Fang
 * Grayor, Wolf Assassin
 * Saviks, Queenbreaker
 * Queenbreaker Captain
 * Queenbreaker Vandal
 * Kaliks-12
 * Wolf High Servitor
 * Peekis, the Disavowed
 * Pirsis, Pallas-Bane
 * Veliniks, the Ravenous
 * Weksis, the Meek
 * Yavek, Wolf Baron
 * Twisted Claw
 * Howling Raiders
 * Wolf Scavengers
 * Wolf Enforcers
 * Wolves' Guard

House of Kings

 * Paskin, King Baron
 * Vekis, King Baron

The Reef

 * Queen Mara Sov
 * Prince Uldren Sov
 * Royal Awoken Guard
 * Petra Venj
 * Variks, the Loyal

The Last City

 * Guardians